Joe Cirilo, a student in the Biomedical Sciences PhD program, is seen at work in a College of Medicine lab in June 2020.

Biomedical Sciences PhD

Training Leaders in Biomedical Research

The Penn State Biomedical Sciences (BMS) PhD Program – with its options in Biochemistry, Genetics, and Genomics, Cancer Biology, Cellular and Integrative Physiology, Translational Therapeutics, and Virology and Immunology – is a nationally and internationally recognized interdisciplinary graduate program that provides students curricular and research training with a unique focus on human health and disease.

Students receive rigorous training that provides the skills necessary to be leaders in biomedical research and other endeavors including business, education, law, journalism, and public policy.

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About the Program

Curriculum

Students in the BMS Graduate Program have the ability to choose among several curricular tracks:

  • Biomedical Sciences
  • Option in Biochemistry, Genetics and Genomics
  • Option in Cancer Biology
  • Option in Cellular and Integrative Physiology
  • Option in Translational Therapeutics
  • Option in Virology and Immunology

The program offers extensive flexibility to tailor curriculum and dissertation research to each individual student’s interests.

With the research emphasis of the program, students also participate in research rotations in at least three laboratories during the first year before choosing the laboratory for their dissertation research.

Learn more about the Biomedical Sciences PhD curriculum.

Get details at an upcoming recruitment event.

Recent Publications by Students

  • Estrogen receptor signaling mechanisms.
    Fuentes N, Silveyra P. Adv Protein Chem Struct Biol. 2019;116:135-170
  • Rivaroxaban for Thromboprophylaxis in High-Risk Ambulatory Patients with Cancer.
    Khorana AA, Soff GA, Kakkar AK, Vadhan-Raj S, Riess H, Wun T, Streiff MB, Garcia DA, Liebman HA, Belani CP, O'Reilly EM, Patel JN, Yimer HA, Wildgoose P, Burton P, Vijapurkar U, Kaul S, Eikelboom J, McBane R, Bauer KA, Kuderer NM, Lyman GH; CASSINI Investigators. N Engl J Med. 2019 Feb 21;380(8):720-728
  • Ezetimibe Added to Statin Therapy after Acute Coronary Syndromes.
    Cannon CP, Blazing MA, Giugliano RP, McCagg A, White JA, Theroux P, Darius H, Lewis BS, Ophuis TO, Jukema JW, De Ferrari GM, Ruzyllo W, De Lucca P, Im K, Bohula EA, Reist C, Wiviott SD, Tershakovec AM, Musliner TA, Braunwald E, Califf RM; IMPROVE-IT Investigators. N Engl J Med. 2015 Jun 18;372(25):2387-97
  • Genome-wide Analyses Identify KIF5A as a Novel ALS Gene.
    Nicolas A, Kenna KP, Renton AE, Ticozzi N, Faghri F, Chia R, Dominov JA, Kenna BJ, Nalls MA, Keagle P, Rivera AM, van Rheenen W, Murphy NA, van Vugt JJFA, Geiger JT, Van der Spek RA, Pliner HA, Shankaracharya, Smith BN, Marangi G, Topp SD, Abramzon Y, Gkazi AS, Eicher JD, Kenna A; ITALSGEN Consortium, Mora G, Calvo A, Mazzini L, Riva N, Mandrioli J, Caponnetto C, Battistini S, Volanti P, La Bella V, Conforti FL, Borghero G, Messina S, Simone IL, Trojsi F, Salvi F, Logullo FO, D'Alfonso S, Corrado L, Capasso M, Ferrucci L; Genomic Translation for ALS Care (GTAC) Consortium, Moreno CAM, Kamalakaran S, Goldstein DB; ALS Sequencing Consortium, Gitler AD, Harris T, Myers RM; NYGC ALS Consortium, Phatnani H, Musunuri RL, Evani US, Abhyankar A, Zody MC; Answer ALS Foundation, Kaye J, Finkbeiner S, Wyman SK, LeNail A, Lima L, Fraenkel E, Svendsen CN, Thompson LM, Van Eyk JE, Berry JD, Miller TM, Kolb SJ, Cudkowicz M, Baxi E; Clinical Research in ALS and Related Disorders for Therapeutic Development (CReATe) Consortium, Benatar M, Taylor JP, Rampersaud E, Wu G, Wuu J; SLAGEN Consortium, Lauria G, Verde F, Fogh I, Tiloca C, Comi GP, Sorarù G, Cereda C; French ALS Consortium, Corcia P, Laaksovirta H, Myllykangas L, Jansson L, Valori M, Ealing J, Hamdalla H, Rollinson S, Pickering-Brown S, Orrell RW, Sidle KC, Malaspina A, Hardy J, Singleton AB, Johnson JO, Arepalli S, Sapp PC, McKenna-Yasek D, Polak M, Asress S, Al-Sarraj S, King A, Troakes C, Vance C, de Belleroche J, Baas F, Ten Asbroek ALMA, Muñoz-Blanco JL, Hernandez DG, Ding J, Gibbs JR, Scholz SW, Floeter MK, Campbell RH, Landi F, Bowser R, Pulst SM, Ravits JM, MacGowan DJL, Kirby J, Pioro EP, Pamphlett R, Broach J, Gerhard G, Dunckley TL, Brady CB, Kowall NW, Troncoso JC, Le Ber I, Mouzat K, Lumbroso S, Heiman-Patterson TD, Kamel F, Van Den Bosch L, Baloh RH, Strom TM, Meitinger T, Shatunov A, Van Eijk KR, de Carvalho M, Kooyman M, Middelkoop B, Moisse M, McLaughlin RL, Van Es MA, Weber M, Boylan KB, Van Blitterswijk M, Rademakers R, Morrison KE, Basak AN, Mora JS, Drory VE, Shaw PJ, Turner MR, Talbot K, Hardiman O, Williams KL, Fifita JA, Nicholson GA, Blair IP, Rouleau GA, Esteban-Pérez J, García-Redondo A, Al-Chalabi A; Project MinE ALS Sequencing Consortium, Rogaeva E, Zinman L, Ostrow LW, Maragakis NJ, Rothstein JD, Simmons Z, Cooper-Knock J, Brice A, Goutman SA, Feldman EL, Gibson SB, Taroni F, Ratti A, Gellera C, Van Damme P, Robberecht W, Fratta P, Sabatelli M, Lunetta C, Ludolph AC, Andersen PM, Weishaupt JH, Camu W, Trojanowski JQ, Van Deerlin VM, Brown RH Jr, van den Berg LH, Veldink JH, Harms MB, Glass JD, Stone DJ, Tienari P, Silani V, Chiò A, Shaw CE, Traynor BJ, Landers JE. Neuron. 2018 Mar 21;97(6):1268-1283.e6

What's New in Biomedical Sciences

More About Penn State College of Medicine

In this video, explore graduate education opportunities at Penn State College of Medicine.

View accessible video with audio description and transcript here.

Student Resources

Penn State College of Medicine strives to create the best culture of respect and inclusion for our students. A number of programs, services and facilities are available.

See more resources for current students

Campus Life

Student Housing

Housing is available to Penn State College of Medicine students, residents and postdocs in University Manor East and West apartment complexes. One of the complexes is pictured as seen from a higher floor of a nearby building, with a road and trees framing it in the background.
Housing in University Manor apartments is available to Penn State College of Medicine students.

Penn State College of Medicine offers housing in its University Manor East or University Manor West apartments to full-time students, residents, postdoctoral scholars and visiting scholars.

Read more about on-campus housing.

Life in Hershey, PA

Penn State College of Medicine is in Hershey, PA, a town known as the home of the HersheyPark amusement park and the Hershey Chocolate Factory. Two banners depicting cartoon candy characters are seen on a light pole in downtown Hershey, PA, in summer 2016, with a brick building with large glass windows in the background.
Hershey, Pa., is known as "The Sweetest Place on Earth."

Penn State College of Medicine's primary campus is located in Hershey, Pa. The area provides numerous diverse and delicious dining experiences, sports, concerts and entertainment, and is a short drive away from the Baltimore, Philadelphia and Washington, DC, metro areas.

Read more about life in Hershey.